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Upon coming aboard Galactica, she is drawn to a meeting held by Brother John Cavil in the chapel he has set up deep within Galactica. Here, Cavil hatches with her a plan to discredit Gaius Baltar and derail his work on the Cylon Detector (TRS: "The Plan"). She fabricates a disc with surveillance footage that seems to indicate that Baltar was responsible for destroying the defense mainframes at the Colonial Ministry of Defense prior to the Fall of the Twelve Colonies and presents it to Commander Adama.
During her time on Galactica, "Godfrey" reveals her underlying nature to be similar to that of her sisters as she attempts to seduce William Adama. She also claims to have had a romantic relationship with the late Dr. Amorak. Her advances make Adama suspicious, and he assigns marines to trail her discretely. When an angry Baltar confronts her in the public head she professes her ignorance of her Cylon nature and maintains the facade of her mission long enough to force him out of her stall and slam the door shut, at which point she breaks down and begins weeping as he rants and raves outside (TRS: "The Plan").
The evidence she presents is initially determined to be authentic, which results in Baltar's arrest. Lieutenant Gaeta later determines that the footage was doctored, much to Baltar's astonishment. Gaeta comments that once the photo was fully resolved, it was almost too easy to find the markers that pointed to photomanipulation - like Godfrey wanted to be found out. Godfrey mysteriously manages to evade the marines and disappear as she is pulled into a hatch by Cavil while "Tough Six" - another copy of Number Six whose cover was as a prostitute - takes her place and escapes by presumably removing her disguise after getting far enough away. Before Godfrey can be captured, Cavil blows her out an airlock above her protests, telling her to inform the other Cylons of the Fleet's location once she resurrects (TRS: "The Plan").
The humans surmise that Godfrey was part of a Cylon plan to discredit Baltar because of his Cylon detector. This "trial by fire" results in a rise in Baltar's popularity within the fleet and protects him against any further charges of complicity in the downfall of the Twelve Colonies. Baltar asks his Messenger Six, who disappeared during Godfrey's presence on Galactica and reappeared when Godfrey vanished, if this was part of the divine plan all along, but she remains enigmatic (TRS: "Six Degrees of Separation").
A file photo of Godfrey exists in President Roslin's office aboard Colonial One as part of a dossier of identified humanoid Cylons. When, where, and how this photo was acquired is not known (TRS: "Epiphanies").